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| 干旱情形下作物叶片气体交换模拟问题及展望 |
| Modeling Leaf Gas Exchange Under Drought: Challenges and Future Prospects |
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| DOI： |
| 中文关键词: 气体交换 叶片尺度 气孔导度模型 干旱 气孔和非气孔因素 |
| 英文关键词:Gas exchange Leaf scale Stomatal conductance model Drought Stomatal and non-stomatal factors |
| 基金项目:新疆维吾尔自治区重大科技专项(2024A03011);国家自然科学基金项目（42230611，42005097）；博士后科研基金项目（BSH2022001）。 |
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| 中文摘要: |
| 研究总结了当前植物叶片气体交换模型的研究进展，重点探讨了气孔导度模型（如BWB模型）和光合生化模型（如FvCB模型）在干旱条件下的局限性。研究发现，干旱会通过气孔和非气孔因素（如光合能力和叶肉导度）影响叶片气体交换，但现有模型在模拟干旱响应时存在不足，无法准确反映水分利用效率（WUE）和内禀水分利用效率（IWUE）的阶段性变化。提出未来研究应通过试验分析模型关键参数（气孔导度斜率m、最大羧化速率Vcmax、最大电子传递速率Jmax和叶肉导度gm）对干旱的响应特征，构建水分响应函数，以改进模型模拟精度。此外，还需明确干旱不同阶段气孔和非气孔因素的调控机制，为理论研究和模型开发提供依据。 |
| 英文摘要: |
| Drought is one of the primary environmental factors affecting gas exchange in crop leaves, and accurately simulating leaf gas exchange under drought conditions is crucial for assessing the impacts of climate change on ecosystems and crop yields. This paper reviews current progress in models of leaf gas exchange in plants, focusing on the limitations of stomatal conductance models (e.g., the BWB model) and photosynthetic biochemical models (e.g., the FvCB model) under drought conditions. Studies have shown that drought affects leaf gas exchange through both stomatal and non-stomatal factors (e.g., photosynthetic capacity and mesophyll conductance), but existing models have limitations in simulating drought responses and cannot accurately reflect the stage-specific changes in water use efficiency (WUE) and intrinsic water use efficiency (IWUE). Future studies should experimentally analyze the response characteristics of key model parameters to drought, including the stomatal conductance slope (m), maximum carboxylation rate (Vcmax), maximum electron transport rate (Jmax), and mesophyll conductance (gm), and develop water-response functions to improve model simulation accuracy. Additionally, the regulatory mechanisms of stomatal and non-stomatal factors at different drought stages need to be clarified to provide a basis for theoretical research and model development. |
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